It’s Thing #16909 from faberdasher, directly enlarged from Thing #12208 by emmett (who is also a central node in the Thingiverse cube gears phenomenon). All of these complex bevel-geared kinetic sculptures can be traced back to the mechanical papercraft of Haruki Nakamura, which was, I believe, first adapted for fused-filament printing in Thing #4683 by Greg Frost.